Page path:
  • Home
  • Julie Meilland

Dr. Julie Meilland


Micropaleontology - Paleoceanography


+49 421 218-65815



UFT 1570

Julie Meilland

Research interests

My research focuses on modern planktonic foraminifera ecology and contribution to the biological carbon pump.

By studying foraminifera assemblages (concentration and taxonomy), morphometry, vertical and horizontal distribution, I aim to better constrain their living habitat and the factors controlling it.

I am currently working within the RECEIVER research theme of the Cluster of Excellence > The Ocean Floor - Earth's Uncharted Interface on the contribution of planktonic foraminifera to the export of carbonate and organic carbon to the deep ocean. The research I carry out in this position is a continuation of my previous work (e.g. own DFG project "Can patchiness explain habitat variability in planktonic foraminifera?" - FoPa).

Icebergs in the Scotia Sea


MARUM Women's representatives (January 2022 - present)

MARUM Postdoc representatives (2019 - 2020)

Falklands penguins
Snow storm onboard the James Clark Ross



  • S. Chaabane, T. de Garidel-Thoron, X. Giraud, J. Meilland, G.-J. Brummer, L. Jonkers, G. Mortyn, M. Greco, N. Casajus, M. Kucera, O. sulpis, A. Kuroyanagi, H. Howa, G. Beaugrand, R. Schiebel, 2024. Size normalizing planktonic Foraminifera abundance in the water column. Limnology and Oceanography Methods, DOI: 10.1002/lom3.10637
  • F. Sykes, J. Meilland, A. Westgård, T. Chalk, M. Chierici, G. Foster, M. Ezat, 2024. Large-scale culturing of the subpolar foraminifera Globigerina bulloides reveals tolerance to a large range of environmental parameters associated to different life-strategies and an extended lifespan. Journal of plankton research, DOI: 10.1093/plankt/fbae029
  • J. Meilland , M. Siccha, R. Morard, M. Kucera, 2024 . Continuous reproduction of planktonic foraminifera in laboratory culture. Journal of Eukaryotic Microbiology , DOI:10.1111/jeu.13022
  • R. Morard, KF Darling, AKM Weiner, C. Hassenrück, C. Vanni, T. Cordier, N. Henry, M. Greco, NM Vollmar, T. Milivojevic, SN Rahman, M. Siccha, J. Meilland , L. Jonkers, F. Quillévéré, G. Escarguel, CJ Douady, T. de Garidel-Thoron, C. de Vargas, M. Kucera, 2024 . The global genetic diversity of planktonic foraminifera reveals the structure of cryptic speciation in plankton. Biological Reviews , DOI:10.1111/brv.13065


  • M. Siccha, R. Morard, J. Meilland , S. Iwasaki, M. Kucera, K. Kimoto, 2023 . Collection of X-ray micro computed tomography images of shells of planktic foraminifera with curated taxonomy. Scientific Data , DOI:10.1038/s41597-023-02498-0
  • G. Anglada-Ortiz, J. Meilland , P. Ziveri, M. Chierici, A. fransson, E. Jones, TL Rasmussen, 2023 . Seasonality of marine calcifies in the northern Barents Sea: Spatiotemporal distribution of planktonic foraminifers and shelled pteropods and their contribution to carbon dynamics. Progress in Oceanography , DOI:10.1016/j.pocean.2023.103121
  • A. Westgård, M. Ezat, TB Chalk, M. Chierici, GL Foster and J. Meilland , 2023 . Large-scale cultivation of Neogloboquadrina pachyderma , its growth in, and tolerance of, variable environmental conditions. Journal of plankton research , DOI:10.1093/plankt/fbad034
  • M. Greco, A. Westgård, F. Sykes, M. Ezat and J. Meilland , 2023 . Uncovering hidden structures: previously hidden pseudopodia and ectoplasmic structures in planktonic foraminifera. Journal of plankton research , DOI:10.1093/plankt/fbad031
  • S. Chaabane, T. de Garidel-Thoron, X. Giraud, R. Schiebel, G. Beaugrand, G.-J. Brummer, N. Casajus, M. Greco, M. Grigoratou, H. Howa, L. Jonkers, M. Kucera, A. Kuroyanagi, J. Meilland , F. Monteiro, G. Mortyn, A. Almogi-Labin, H. Asahi, S. Avnaim-Katav, F. Bassinot, CV Davis, DB Field, I. Hernández-Almeida, B. Herut, G. Hosie, W. Howard, A. Jentzen, D. G. Johns, L. Keigwin, J. Kitchener, KE Kohfeld, DVO Lessa, C. Manno, M. Marchant, S. Ofstad, JD Ortiz, A. Post, A. Rigual-Hernandez, MC Rillo, K. Robinson, T. Sagawa, F. Sierro, KT Takahashi, A. Torfstein, I. Venancio, M. Yamasaki and P. Ziveri, 2023 . The FORCIS database: A global census of planktonic Foraminifera from ocean waters. Scientific Data , DOI:10.1038/s41597-023-02264-2
  • C. Hayatte Akhoudas, J.-B. Sallée, G. Reverdin, FA Haumann, E. Pauthenet, CC Chapman, F. Margirier, C. Lo Monaco, N. Metzl, J. Meilland , C. Stranne, 2023 . Isotopic evidence for an intensified hydrological cycle in the Indian sector of the Southern Ocean. Nature communications , DOI:10.1038/s41467-023-38425-5
  • J. Meilland , M. Ezat, A. Westgård, C. Manno, R. Morard, M. Siccha, M. Kucera, 2023 . Rare but persistent asexual reproduction explains the success of planktonic foraminifera in polar oceans. Journal of Plankton Research , DOI:10.1093/plankt/fbac069


  • F. Tell, L. Jonkers, J. Meilland , M. Kucera, 2022 . Upper-ocean flux of biogenic calcite produced by the Arctic planktonic foraminifera Neogloboquadrina pachyderma . Biogeosciences , DOI:10.5194/bg-19-4903-2022
  • T. de Garidel-Thoron, S. Chaabane, X. Giraud, J. Meilland , L. Jonkers, M. Kucera, GJ. A. Brummer, M. Grigoratou, FM Monteiro, M. Greco, PG Mortyn, A. Kuroyanagi, H. Howa, G. Beaugrand, R. Schiebel, 2022 . The Foraminiferal Response to Climate Stressors Project: Tracking the Community Response of Planktonic Foraminifera to Historical Climate Change. Frontiers in Marine Science , DOI:10.3389/fmars.2022.827962
  • L. Jonkers, GJA, Brummer, J. Meilland , J. Groeneveld, M. Kucera, 2022 . Variability in Neogloboquadrina pachyderma stable isotope ratios from isothermal conditions: implications for individual foraminifera analysis. Climate of the Past , DOI:10.5194/cp-18-89-2022
  • J. Meilland , P. Cornuault, R. Morard, GJA, Brummer, M. Kucera, 2022 . Identification guide to extant planktonic foraminifera. Part 1: Family Candeinidae and genera Berggrenia , Bolivina , Dentigloborotalia , and Neogallitellia . ICES Identification Leaflets for Plankton No. 196, DOI:10.17895/


  • J. Meilland , M. Siccha, M. Kaffenberger, J. Bijma, M. Kucera, 2021 . Population dynamics and reproduction strategies of planktonic foraminifera in the open ocean. Biogeosciences , DOI:10.5194/bg-18-5789-2021
  • K. Zamelczyk, A. Fransson, M. Chierici, E. Jones, J. Meilland , G. Anglada-Ortiz, H. Lødemel, 2021 . Distribution and Abundances of Planktic Foraminifera and Shelled Pteropods During the Polar Night in the Sea-Ice Covered Northern Barents Sea. Frontiers in Marine Sciences , DOI:10.3389/fmars.2021.644094
  • L. Jonkers, J. Meilland , MC Rillo, T. de Garidel-Thoron, JA Kitchener, M. Kucera, 2021 . Linking zooplankton time series to the fossil record. ICES Journal of marine sciences , DOI: 10.1093/icesjms/fsab123
  • G. Anglada-Ortiz, K. Zamelczyk, J. Meilland , P. Ziveri, M. Chierici, A. Fransson, TL Rasmussen, 2021 . Planktic foraminiferal and pteropod contributions to carbon dynamics in the Arctic Ocean (north Svalbard margin). Frontiers in Marine Sciences , DOI:10.3389/fmars.2021.661158
  • KAF Zonneveld, J. Meilland , B. Donner, GJM Versteegh, 2021 . Export flux succession of dinoflagellate cysts and planktonic foraminifera in an active upwelling cell off Cape Blanc (NW Africa). European Journal of Phycology, DOI:10.1080/09670262.2021.1885066


  • M. Greco, J. Meilland , K. Zamelczyk, TL Rasmussen, M. Kucera, 2020 . The effect of an experimental decrease in salinity on the viability of the Subarctic planktonic foraminifera Neogloboquadrina incompta. Polar Research , DOI:10.33265/polar.v39_3842
  • J. Meilland , H. Howa, V. Hulot, I. Demangel, J. Salaün, T. Garlan, 2020 . Population dynamics of modern planktonic foraminifera in the western Barents Sea. Biogeosciences , DOI: 10.5194/bg-17-1437-2020
  • S. Ofstad, J. Meilland , K. Zamelczyk, M. Chierici, A. Fransson, F. Gründger, TL Rasmussen, 2020 . Development, productivity and seasonality of living planktonic foraminiferal faunas and Limacina helicina in an area of ​​intense methane seepage in the Barents Sea. JGR Biogeoscienc es , DOI: 10.1029/2019JG005387


  • AY Hsiang, A. Brombacher, MC Rillo, MJ Mleneck-Vautravers, S. Conn, S. Lordsmith, A. Jentzen, MJ Henehan, B. Metcalfe, I. Fentpn, BS Wade, L. Fox, J. Meilland , CV Davis, U. Baranowski, J. Groeneveld, KM Edgar, A. Movellan, T. Aze, HJ Dowsett, CG Miller, N. Rios, PM Hull, 2019 . Endless Forams: >34,000 modern planktonic foraminiferal images for taxonomic training and automated species recognition using convolutional neural networks. Paleoceanography and Paleoclimatology , DOI: 10.1029/2019PA003612.
  • J. Meilland , M. Siccha, M. Weinkauf, L. Jonkers, R. Morard, U. Baranowski, A. Baumeister, J. Bertlich, GJ Brummer, P. Debray, T. Fritz-Endres, J. Groeneveld, L Magerl, P. Munz, M. Rillo, C. Scmidt, H. Takagi, G. Theara, M. Kucera, 2019 . Highly replicated sampling reveals no vertical migration but stable species-specific vertical habitats in planktonic foraminifera. Journal of Plankton Research , (41) 2, 127-14.


  • R. Schiebel, SM Smart , A. Jentzen, L. Jonkers, R. Morard, J. Meilland , E. Michel, H. Coxall, P. Hull, T. de Garidel-Thoron, T. Aze, F. Quillevere, H. Ren, D. Sigman, HB Vonhof, A. Martinez-Garcia, M. Kucera, J. Bijma, H. Spero, GH Haug, 2018 . Advances in planktonic foraminifer research: New perspectives for paleoceanography. Revue de Micropaleontologie, 61, 113-138 .
  • A. André, F. Quillévéré, R. Schiebel, R. Morard, H. Howa, J. Meilland , C. Douady, 2018 . Disconnection between genetic and morphological diversity in the planktonic foraminifer Neogloboquadrina pachyderma from the Indian Sector of the Southern Ocean , Marine Micropaleontology , 144, 14-24.
  • J. Meilland , R. Schiebel, C. Lo Monaco, S. Sanchez, H. Howa, 2018 . Abundances and tests for planktic foraminifera across the Southwest Indian Ocean: Implications for carbon fluxes. Deep Sea Research I , 131, 27-47.


  • R. Schiebel, R. Spielhagen, J. Garnier, J. Hagemann, H. Howa, A. Martinez-Garcia, J. Meilland , E. Michel, I. Salter, M. Yamasaki, G. Haug, 2017 . Modern planktic foraminifera in the high-latitude ocean. Marine Micropaleontology , 136, 1-13.


  • J. Meilland , H. Howa, C. Lo Monaco, R. Schiebel, 2016 . Individual planktic foraminifera protein biomass affected by trophic conditions in the Southern Indian Ocean [30°S-60°S]. Marine Micropaleontology , 124, 63-74.
  • M. Reambeauville, J. Meilland , P. Ziveri, R. Schiebel, S. Blain, I. Salter, 2016 . Planktic foraminifera and coccoliths contribute to carbonate fluxes over the central Kerguelen plateau. Deep Sea Research I , 111, 91-101.
  • J. Meilland , S. Fabri-Ruiz, P. Koubbi, C. Lo Monaco, C. Cotte, G. Hosie, S. Sanchez, H. Howa, 2016 . Planktonic foraminiferal biogeography in the Southern Indian Ocean: Contribution from CPR data. Deep Sea Research I , 110, 75-89.


  • L. Subirana, B. Péquin, S. Michely, ML Escande, J. Meilland , E. Derelle, G. Piganeau, Y. Desdevises, H. Moreau, N. Grimsley, 2013 . Morphological variations and general plasticity in cryptic species of Eastern reococcus (Mameillophyceae, Mamiellales) isolated from the Mediterranean sea. Protist , Vol. 164, 643-659.

Research Expeditions

09-10 / 2022: RV Maria S. Merian, MSM111 (Iceland - Baffin Bay - Canada), 5 weeks

06 / 2022: Co-chief scientist, CAGE 22-4 cruise within the ARCLIM project and with UiT, the Arctic University of Norway (Nordic Seas), RV Helmer Hanssen, 1 week

07 / 2021: Co-chief scientist, CAGE 21-2 cruise within the ARCLIM project and with UiT, the Arctic University of Norway (Fram Strait), RV Helmer Hanssen, 1 week

08-09 / 2020: RV Celtic Explorer, expedition CIAAN with the University of Galway (Nordic Seas), 4 weeks

11-12 / 2019: RSS James Clark Ross, expedition JR19001 with the British Antarctic Survey (Antarctica - Scotia Sea), 7 weeks

10 / 2018: Co-chief scientist, RV Helmer Hanssen, teaching cruise (Norwegian coast), 3 days

08-09 / 2017: RV Meteor, M-140 FORAMFLUX (Cabo Verde - Canary Island), 4 weeks

10 / 2016: Co-chief scientist, RV Helmer Hanssen, teaching cruise (Norwegian coast), 1 week

07 / 2016: RV Atalante, IFREMER, STEP cruise (Storfjorden), 3 weeks

06 / 2016: RV Helmer Hanssen, CAGE-ROV cruise (Svalbard - Barents Sea), 3 weeks

04 / 2016: RV Helmer Hanssen, CAGE-16 expedition (Savlbard), 1 week

04 / 2016: RV Helmer Hanssen, CAGE-15 expedition (Bjørnøyrenna), 1 week

02 / 2016: RV Helmer Hanssen, CAGE-OA expedition (Svalbard to Tromsø), 2 weeks

08-09 / 2014: RV Pourquoi Pas ?, IFREMER, expedition MOCOSED (N-Atlantic, Barents Sea), 6 weeks

01-03 / 2014: RV Marion Dufresne, IPEV, expedition MD197 - MYCTO (Southern Indian Ocean), 7 weeks

01-03 / 2013: RV Marion Dufresne, IPEV, expedition MD193 - LOGIPEV (Southern Indian Ocean), 7 weeks

01-03 / 2012: RV Marion Dufresne, IPEV, expedition MD189 - India South 2 (Southern Indian Ocean), 7 weeks

01-03 / 2011: RV Marion Dufresne, IPEV, MD185 expedition - India Southern (Southern Indian Ocean), 8 weeks